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NRF Connect Programmer bootloader signature error when not flashing bootloader

Similar to https://devzone.nordicsemi.com/f/nordic-q-a/61935/bootloader-flashing-error---nordic-nrf52840-usb-dongle I get the following error:

DFU failed: Error message for known extended error code from DFU target: The init packet does not contain a signature. This bootloader requires DFU updates to be signed.
However, I'm not trying to update the bootloader, I'm trying to upload an application (the NRF sniffer .hex) onto an nrf28540 dongle (Specifically Laird Connectivity BL654, but that should be irrelevant?), as can be seen on the following image:

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